Cordia bicolor    DC.

Occasional evergreen or briefly deciduous tree (5-8 m) found in areas offering abundant sunlight and characterized by secondary vegetation. Along with Balsa, Juco, and Cecropia, this species is a principle participant in the forest's struggle to reclaim disturbed land.

Description: A fast-growing, short-lived "pioneer" species, Cordia has a cylindrical trunk (10 cm) that often becomes inclined as it grows rapidly toward the brightest source of light available. The tree is clad in smooth and silvery-gray bark. Possessing only a few, long, drooping branches, the crown of this species is made moderately dense by the large size of its foliage. Leaves are simple, alternate, and arranged in two tiers along the twigs. Though generally large, they are quite variable in shape and size. On average, the leaves are elliptical in form and measure 20 cm (16-23 cm) long by 7 cm (6-9 cm) wide. All leaf surfaces, twigs, and petioles (1 cm) are covered with short, stiff hairs that give them a rough, sandpapery texture. Annually, leaves slowly senesce and are shed during the course of the dry season (December-April). By late April, when trees are completely bare, a new growth spurt begins and the Cordia crowns are renewed by May. The white flowers (8 mm long by 5 mm in diameter) appear soon thereafter in large terminal panicles. They each consist of a five pointed, green calyx; five, backwardly curving white petals; five long, forward-pointing stamens; and a central, three-part pistil. The timing of flowering periods is variable from year to year and between individuals, but trees are fertile sometime in May, June, and/or July. Glossy, green, globular fruits (1.5 cm), set in the same - now wider - calyxes, mature as they pale to dull yellow or white and become translucent. In them, a solitary seed is surrounded by a thick layer of slippery, jelly-like pulp. Harvests are annual and occur in September and October.

Similar Species: The large size of Cordia bicolor foliage, coupled with its nature as a small, secondary species found in areas of rapid regeneration, aid in its identification.

Natural History: Cordia flowers are pollinated by wasps and butterflies, seen visiting them in great numbers during periods of peak fertility. Bats and white-faced monkeys play a role in dispersing Cordia fruit (Croat, 1978). Cordia is one of this ecosystem's most important secondary species, able to support the harsh, adverse conditions created when the primary forest tree-cover is damaged or destroyed. Its ability to grow, even thrive, under such extreme circumstances is important to the process of forest succession. Cordia's rapid growth quickly establishes an anchoring root system as well as a sheltering crown that help to protect exposed soils and provide the more consistent regimes of humidity and temperature that are required by most other plant species. The early establishment of a pioneer plant community after environmental damage has taken place is one of the critical first steps nature implements in its effort to jump-start the regeneration of the primary forest.

Uses: Cordia bicolor could be used in large-scale reforestation efforts. The tree, planted along with other pioneer species, produces a non-herbaceous ground cover that protects and enriches the soil with organic matter and that provides shade, increases humidity, and moderates temperatures.

Distribution: This Cordia may be found in very humid forests wherever sunlight reaches the ground in abundance. Such areas include secondary forests as well as road, trail, and streamsides. Cordia is native to the wet or humid parts of both of Costa Rica's lowland slopes, being absent only in drier Guanacaste. It ranges from Mexico and the Antilles to northern South America (Zamora, 1989).